Ecology — Hard Practice Quiz
A Biology cheat sheet for Ecology — every key formula with its symbols defined — plus a hard-level practice quiz to test recall.
Formulas & key concepts
<b>Ecology</b>: Study of interactions among organisms and between organisms and their environment. <b>Biosphere</b>: All life on Earth + all parts where life exists (land, water, atmosphere, 8km above to 11km below ocean surface).
<b>Individual organism</b> → <b>Population</b> (same species, same area) → <b>Community</b> (different populations in area) → <b>Ecosystem</b> (community + physical environment) → <b>Biome</b> (similar climates, typical organisms) → <b>Biosphere</b>.
<b>Biotic</b>: Living components (animals, plants, bacteria, fungi). <b>Abiotic</b>: Non-living (sunlight, temperature, water, precipitation, soil, wind, humidity). Interact to shape ecosystem.
<b>Observation</b>: Identify species, behaviors, patterns. <b>Experimentation</b>: Test hypotheses, manipulate variables (greenhouse studies, field experiments). <b>Modeling</b>: Predict long-term/large-scale events, based on data.
<b>Habitat</b>: Physical location where organism lives (address). <b>Niche</b>: Organism's role in ecosystem (profession) - what it eats, when active, where lives, how interacts. <b>Fundamental niche</b> (potential) vs. <b>Realized niche</b> (actual with competition).
<b>Autotrophs</b>: Make own food. <b>Photosynthesis</b>: Use sunlight (plants, algae, cyanobacteria: 6CO₂ + 6H₂O + light → C₆H₁₂O₆ + 6O₂). <b>Chemosynthesis</b>: Use chemical energy (deep-sea vent bacteria, sulfur compounds). Foundation of food chains.
<b>Herbivores</b>: Eat plants (primary consumers). <b>Carnivores</b>: Eat animals (secondary/tertiary consumers). <b>Omnivores</b>: Eat both plants + animals. <b>Scavengers</b>: Eat dead animals. <b>Detritivores</b>: Eat detritus (earthworms, crabs). <b>Decomposers</b>: Break down organic matter (bacteria, fungi).
Linear sequence showing energy transfer. Example: Grass (producer) → Rabbit (primary consumer/herbivore) → Fox (secondary consumer/carnivore) → Decomposers. Each step = <b>trophic level</b>.
Complex network of interconnected food chains. Shows multiple feeding relationships. More realistic than simple chains. Organisms often eat multiple things and are eaten by multiple predators.
<b>10% Rule</b>: Only ~10% of energy transfers to next trophic level (90% lost as heat from metabolism). <b>Biomass pyramid</b>: Total mass of living tissue decreases at higher levels. Limits food chain length to ~4-5 levels.
Efficiency of energy transfer between levels. Typically 5-20% (average 10%). Lost energy: respiration (heat), waste, incomplete digestion. Why top predators are rare.
<b>Decomposers</b> (bacteria, fungi): Break down dead organisms, return nutrients to soil. Essential for recycling. <b>Detritus</b>: Dead organic matter. Without decomposers, nutrients locked up in dead bodies.
Energy from chemical reactions (not sunlight). <b>Deep-sea vents</b>: Bacteria use H₂S + O₂ → organic compounds. Support entire ecosystems without photosynthesis. Also in hot springs, tidal marshes.
<b>Evaporation</b> (liquid → vapor), <b>Transpiration</b> (plants release water), <b>Condensation</b> (vapor → clouds), <b>Precipitation</b> (rain/snow), <b>Runoff</b> (flows to rivers/oceans), <b>Infiltration</b> (groundwater). Driven by solar energy.
<b>Photosynthesis</b>: CO₂ → organic compounds. <b>Respiration</b>: Organic → CO₂ (returns to atmosphere). <b>Decomposition</b>: Dead organisms → CO₂. <b>Combustion</b>: Burning fossil fuels/wood → CO₂. <b>Oceans</b>: Dissolve/release CO₂. <b>Sediments</b>: Long-term storage (fossil fuels).
<b>Nitrogen fixation</b>: N₂ → NH₃/NH₄⁺ (bacteria: Rhizobium in root nodules, cyanobacteria, lightning). <b>Nitrification</b>: NH₄⁺ → NO₂⁻ → NO₃⁻ (bacteria). <b>Assimilation</b>: Plants absorb NO₃⁻, make proteins. <b>Ammonification</b>: Dead organisms → NH₄⁺. <b>Denitrification</b>: NO₃⁻ → N₂ (back to atmosphere).
<b>Weathering</b>: Rocks release phosphate (PO₄³⁻). <b>Absorption</b>: Plants take up from soil. <b>Food chain</b>: Passes through consumers. <b>Decomposition</b>: Returns to soil. <b>Sedimentation</b>: Washes to ocean, forms rocks. <b>No atmospheric component</b> (unlike C, N). Slow cycle.
<b>Law of the Minimum</b>: Nutrient in shortest supply limits growth. <b>Terrestrial</b>: Often nitrogen. <b>Aquatic</b>: Often phosphorus or nitrogen. Adding limiting nutrient increases productivity (can cause eutrophication).
<b>Size</b>: Number of individuals. <b>Density</b>: Number per unit area. <b>Distribution</b>: <b>Clumped</b> (groups, most common), <b>Uniform</b> (evenly spaced, territorial), <b>Random</b> (unpredictable, rare). <b>Age structure</b>: Proportion in each age group.
Population change = (Birth rate + Immigration) - (Death rate + Emigration). <b>Immigration</b>: Individuals moving in. <b>Emigration</b>: Individuals moving out. <b>Demography</b>: Study of population statistics.
<b>J-curve</b>: Accelerating growth when resources unlimited. <b>Equation</b>: dN/dt = rN (r = intrinsic rate of increase). Population doubles at constant intervals. Unsustainable long-term. Occurs after colonization or low density.
<b>S-curve</b>: Growth slows as approaches carrying capacity. <b>Carrying capacity (K)</b>: Maximum population environment can sustain. Growth rate highest at K/2. More realistic than exponential. <b>Equation</b>: dN/dt = rN(K-N)/K.
Maximum population size environment can support indefinitely. Determined by limiting resources (food, water, space, shelter). Varies with environmental conditions. Population fluctuates around K.
Effects depend on population density. <b>Competition</b> (for resources), <b>Predation</b> (more prey = more predators), <b>Disease/Parasitism</b> (spreads faster when crowded), <b>Stress</b>, <b>Waste accumulation</b>. Regulate populations near carrying capacity.
Effects regardless of density. <b>Weather</b> (drought, freeze, flood), <b>Natural disasters</b> (fires, hurricanes), <b>Seasonal cycles</b>, <b>Climate</b>. Can cause drastic population changes.
<b>r-selected</b>: Unstable environments. Many offspring, small size, little parental care, mature quickly, short lifespan (insects, mice, weeds). <b>K-selected</b>: Stable environments. Few offspring, large size, much parental care, mature slowly, long lifespan (elephants, whales, humans).
<b>Type I</b>: Low infant mortality, most die old (humans, large mammals, high parental care). <b>Type II</b>: Constant death rate throughout life (birds, rodents). <b>Type III</b>: High infant mortality, few reach adulthood (fish, insects, plants, many offspring).
<b>Negative feedback</b>: Density-dependent factors slow growth as population increases. <b>Oscillations</b>: Predator-prey cycles (e.g., lynx-hare). <b>Zero population growth</b>: Birth rate = death rate. Populations stabilize around carrying capacity.
<b>Competition</b> (-/-), <b>Predation</b> (+/-), <b>Herbivory</b> (+/-), <b>Mutualism</b> (+/+), <b>Commensalism</b> (+/0), <b>Parasitism</b> (+/-). Shape community structure and evolution.
<b>Interspecific</b>: Between different species for same resources. <b>Intraspecific</b>: Within same species (usually more intense). <b>Competitive exclusion principle</b>: Two species cannot coexist if they occupy identical niche. <b>Resource partitioning</b>: Species divide resources (different times, places, foods).
<b>Predation</b>: One organism kills/eats another. Drives evolution. <b>Prey defenses</b>: <b>Camouflage</b> (blend in), <b>Warning coloration</b> (bright = dangerous), <b>Mimicry</b> (Batesian: harmless looks dangerous; Müllerian: both dangerous look similar), <b>Physical</b> (spines, shells).
Close, long-term interactions. <b>Mutualism (+/+)</b>: Both benefit (bees + flowers, lichen, mycorrhizae). <b>Commensalism (+/0)</b>: One benefits, other unaffected (barnacles on whales). <b>Parasitism (+/-)</b>: Parasite benefits, host harmed (tapeworms, ticks, mistletoe).
Species with disproportionately large impact on ecosystem relative to abundance. Removal causes major changes. Examples: Sea otters (control urchins), wolves (control herbivores), sea stars (control mussels). Maintain biodiversity.
Form major structural component of habitat. Create environment for other species. Examples: Coral (reefs), kelp (forests), trees (forest canopy). Usually abundant, high biomass.
<b>Primary succession</b>: Starts on bare rock/new land (glacial retreat, volcanic island). <b>Pioneer species</b>: Lichens, mosses. <b>Secondary succession</b>: After disturbance (fire, farming, logging), soil present. Faster than primary. <b>Climax community</b>: Stable end point, characteristic of climate.
<b>Species richness</b>: Number of different species. <b>Relative abundance</b>: Evenness of distribution. High biodiversity increases: ecosystem stability, productivity, resilience to disturbance. Highest in tropical rainforests, coral reefs.
<b>Biotic</b>: Producers, consumers (primary, secondary, tertiary), decomposers. <b>Abiotic</b>: Sunlight, temperature, water, soil, minerals, climate. Energy flows through (not recycled), matter cycles.
<b>Gross Primary Productivity (GPP)</b>: Total energy captured by producers. <b>Net Primary Productivity (NPP)</b>: GPP - respiration (energy available to consumers). <b>Standing crop</b>: Total biomass at given time. Highest in tropical rainforests, estuaries, coral reefs.
Determined by temperature + precipitation. <b>Tropical rainforest</b> (high temp, high rain, high diversity). <b>Temperate forest</b> (moderate temp, seasonal, deciduous trees). <b>Grassland</b> (moderate rain, periodic fire). <b>Desert</b> (low rain, extreme temps). <b>Taiga</b> (cold, conifers). <b>Tundra</b> (very cold, permafrost, low diversity).
<b>Freshwater</b>: Lakes (littoral, limnetic, profundal zones), rivers/streams, wetlands (filter water, biodiversity). <b>Marine</b>: <b>Photic zone</b> (sunlit, 0-200m), <b>Aphotic zone</b> (dark). <b>Intertidal</b> (tides), <b>Coral reefs</b> (high diversity), <b>Estuaries</b> (salt + fresh mix, high productivity), <b>Open ocean</b>.
Climate (long-term temperature + precipitation patterns) determines biome. <b>Latitude</b>: Tropical (equator) → temperate → polar. <b>Altitude</b>: Similar to latitude (higher = colder). <b>Rain shadow</b>: Mountains block moisture, desert on leeward side.
<b>Aquatic</b>: <b>Littoral</b> (shallow, near shore), <b>Limnetic</b> (open water, sunlit), <b>Profundal</b> (deep, no light), <b>Benthic</b> (bottom). <b>Terrestrial</b>: <b>Canopy</b> (treetops), <b>Understory</b> (shrubs/small trees), <b>Forest floor</b> (decomposition).
<b>Deforestation</b>: Clearing forests (agriculture, logging, development). <b>Urbanization</b>: Cities replace natural habitats. <b>Fragmentation</b>: Habitats broken into small patches (edge effects, reduced population sizes). Leading cause of species extinction.
<b>Air</b>: Smog, particulates, greenhouse gases (CO₂, CH₄), acid rain precursors (SO₂, NOₓ). <b>Water</b>: Sewage, chemicals, oil spills, thermal pollution, eutrophication. <b>Soil</b>: Pesticides, heavy metals, industrial waste. <b>Noise/Light</b>: Affects animal behavior.
<b>Greenhouse effect</b>: CO₂, CH₄, N₂O trap heat. <b>Global warming</b>: Average temp rising (~1°C since 1880). <b>Impacts</b>: Ice melting, sea level rise, extreme weather, range shifts, coral bleaching, extinction risk. <b>Causes</b>: Fossil fuel burning, deforestation.
Non-native species introduced to new area. Often lack natural predators/parasites. <b>Impacts</b>: Outcompete natives, alter habitats, disrupt food webs, cause extinctions. Examples: Zebra mussels, kudzu, cane toads, Asian carp. Prevention: biosecurity, quarantine.
Harvesting organisms faster than they can reproduce. <b>Overfishing</b>: Cod, tuna population collapse. <b>Overhunting</b>: Passenger pigeon (extinct), whales (endangered). <b>Poaching</b>: Elephants (ivory), rhinos (horns). Unsustainable resource use.
<b>Cultural eutrophication</b>: Excess nutrients (N, P from fertilizer runoff, sewage) enter water. <b>Process</b>: Algal bloom → blocks sunlight → algae die → decomposition uses O₂ → <b>hypoxia/dead zones</b> → fish/organisms die. Major problem in lakes, coastal areas.
<b>Bioaccumulation</b>: Toxins build up in organism over time. <b>Biomagnification</b>: Toxin concentration increases up food chain. Top predators most affected. Examples: DDT (egg shell thinning in eagles), Mercury (fish consumption warnings), PCBs. Fat-soluble toxins persist.
<b>Stratospheric ozone (O₃)</b>: Blocks UV radiation. <b>CFCs</b> (chlorofluorocarbons from refrigerants, aerosols) break down ozone. <b>Ozone hole</b>: Over Antarctica. <b>Effects</b>: Increased UV → skin cancer, cataracts, crop damage. <b>Montreal Protocol</b>: Banned CFCs, ozone recovering.
<b>Causes</b>: SO₂ + NOₓ from fossil fuel combustion → H₂SO₄, HNO₃ in atmosphere. pH < 5.6. <b>Effects</b>: Damages forests (leaches nutrients from soil), acidifies lakes (kills fish/amphibians), corrodes buildings. <b>Solutions</b>: Scrubbers on smokestacks, catalytic converters, renewable energy.
<b>Soil erosion</b>: Tree roots hold soil, removal causes erosion. <b>Species loss</b>: Habitat destruction, extinction. <b>Climate</b>: Less CO₂ absorption, more CO₂ release (burning/decomposition). <b>Water cycle disruption</b>: Less transpiration, altered rainfall. <b>Indigenous displacement</b>.
Science of protecting biodiversity. <b>Goals</b>: Prevent extinctions, preserve habitats, restore ecosystems. <b>Strategies</b>: Protected areas, captive breeding, reintroduction programs, habitat corridors, legislation (Endangered Species Act, CITES).
<b>Renewable</b>: Replace at/faster than use rate (solar, wind, forests, fisheries if managed). <b>Non-renewable</b>: Finite supplies (fossil fuels, minerals). <b>Ecological footprint</b>: Land/water needed to support lifestyle. <b>Sustainability</b>: Meeting needs without compromising future generations.
Active restoration of degraded ecosystems. <b>Examples</b>: Wetland restoration, reforestation, stream restoration, removing invasive species. <b>Reintroduction</b>: Return species to former range (wolves to Yellowstone, condors). <b>Bioremediation</b>: Use organisms to clean pollution.
Areas with high species richness + endemism (species found nowhere else) + threat level. <b>Criteria</b>: >1,500 endemic plant species, >70% habitat loss. <b>Examples</b>: Madagascar, tropical Andes, Caribbean, Philippines. Conservation priorities. ~36 hotspots globally.
<b>National parks</b>, <b>Wildlife reserves</b>, <b>Marine protected areas (MPAs)</b>, <b>Wilderness areas</b>. Preserve habitats, prevent development. <b>Challenges</b>: Insufficient size, isolated (need corridors), enforcement, human-wildlife conflict. ~15% of land, ~7% of ocean protected.
Human population ~8 billion, growing. <b>Demographic transition</b>: High birth/death → high birth/low death (rapid growth) → low birth/death (stable). <b>Carrying capacity</b>: Uncertain for humans (technology extends but finite). <b>Resource consumption</b>: Developed nations use disproportionate share. Need sustainable practices.
Practice quiz
An ecosystem's primary producers collectively capture $10,000 \text{ kcal}$ of energy. Assuming a typical trophic efficiency, how much energy would be available to a tertiary consumer in this food chain, and what is the primary reason for the significant energy loss at each transfer?
- A) $10 \text{ kcal}$; Energy is lost as heat during metabolic processes and incomplete digestion.
- B) $100 \text{ kcal}$; Energy is consumed by decomposers before reaching higher trophic levels.
- C) $1,000 \text{ kcal}$; Most energy is stored in biomass and not transferred.
- D) $1 \text{ kcal}$; Energy is primarily used for reproduction at lower trophic levels.
Answer: A) $10 \text{ kcal}$; Energy is lost as heat during metabolic processes and incomplete digestion.
A population of deer is introduced to a new, resource-rich island. Initially, its growth follows a J-curve, but as the population increases, it transitions to an S-curve. At what point in the S-curve is the population's growth rate highest, and which factors are primarily responsible for the eventual stabilization of the population around the island's carrying capacity?
- A) When the population size is close to the carrying capacity ($K$); Density-independent factors like severe weather.
- B) When the population size is half of the carrying capacity ($K/2$); Density-dependent factors such as competition for food and disease.
- C) When the population size is very small; Density-independent factors like natural disasters.
- D) When the population size slightly exceeds the carrying capacity ($K$); Density-dependent factors like increased predation.
Answer: B) When the population size is half of the carrying capacity ($K/2$); Density-dependent factors such as competition for food and disease.
Two species of closely related birds, Species A and Species B, feed on the same type of insect in the same forest. However, Species A primarily forages in the upper canopy, while Species B forages in the understory. If a new invasive predator significantly reduces the population of Species B, allowing Species A to expand its foraging range to include the understory, which ecological principles are best illustrated by the initial coexistence and the subsequent expansion of Species A?
- A) Initial coexistence by competitive exclusion; Expansion due to mutualism.
- B) Initial coexistence by resource partitioning; Expansion due to a shift from realized niche to fundamental niche.
- C) Initial coexistence by commensalism; Expansion due to increased carrying capacity.
- D) Initial coexistence by predation; Expansion due to reduced density-independent factors.
Answer: B) Initial coexistence by resource partitioning; Expansion due to a shift from realized niche to fundamental niche.
Following a massive volcanic eruption that completely sterilizes a large land area, creating new rock formations, what type of ecological succession would occur, and what role do the initial colonizing organisms play in preparing the environment for later successional stages?
- A) Secondary succession; Pioneer species like grasses quickly establish, adding organic matter to existing soil.
- B) Primary succession; Pioneer species such as lichens and mosses break down rock and contribute to soil formation.
- C) Primary succession; K-selected species immediately dominate, stabilizing the ecosystem.
- D) Secondary succession; Decomposers rapidly break down volcanic ash, releasing nutrients for plant growth.
Answer: B) Primary succession; Pioneer species such as lichens and mosses break down rock and contribute to soil formation.
Deforestation is a major contributor to increased atmospheric $CO_2$ levels, exacerbating climate change. Describe two distinct mechanisms by which large-scale deforestation impacts the carbon cycle, linking these to fundamental biological processes.
- A) Reduced nitrogen fixation by trees and increased methane release from exposed soil.
- B) Decreased photosynthesis, leading to less $CO_2$ uptake, and increased $CO_2$ release from the decomposition or burning of cleared biomass.
- C) Enhanced transpiration, which cools the atmosphere, and increased carbon sequestration in new agricultural crops.
- D) Accelerated water cycle, leading to more precipitation, and reduced respiration rates in remaining vegetation.
Answer: B) Decreased photosynthesis, leading to less $CO_2$ uptake, and increased $CO_2$ release from the decomposition or burning of cleared biomass.
Nitrogen is often a limiting nutrient in terrestrial ecosystems. Explain how two distinct bacterial processes contribute to making atmospheric nitrogen available to plants and then returning it to the atmosphere, and how human activities can significantly alter the natural balance of these processes.
- A) Nitrification converts $N_2$ to $NH_4^+$, and ammonification converts $NO_3^-$ to $N_2$; Human use of pesticides inhibits these bacteria.
- B) Nitrogen fixation converts $N_2$ to $NH_3/NH_4^+$, and denitrification converts $NO_3^-$ to $N_2$; Human activities like fertilizer use and fossil fuel combustion add excess reactive nitrogen to ecosystems.
- C) Assimilation converts $N_2$ to organic compounds, and decomposition converts organic compounds to $N_2$; Human deforestation reduces plant assimilation.
- D) Weathering releases $N_2$ from rocks, and runoff carries it to oceans; Human mining of nitrogen-rich rocks accelerates this.
Answer: B) Nitrogen fixation converts $N_2$ to $NH_3/NH_4^+$, and denitrification converts $NO_3^-$ to $N_2$; Human activities like fertilizer use and fossil fuel combustion add excess reactive nitrogen to ecosystems.
A persistent, fat-soluble toxin is introduced into an aquatic ecosystem. If primary producers in this ecosystem accumulate the toxin at a concentration of $0.005 \text{ ppm}$, what would be the approximate concentration of this toxin in a tertiary consumer, and why are top predators particularly susceptible to the harmful effects of such toxins?
- A) $0.05 \text{ ppm}$; Top predators have faster metabolic rates, leading to quicker toxin breakdown.
- B) $0.5 \text{ ppm}$; Top predators consume a wider variety of food sources, diluting the toxin.
- C) $5 \text{ ppm}$; Top predators accumulate toxins through biomagnification, as they consume large quantities of contaminated prey over their lifespan.
- D) $0.0005 \text{ ppm}$; Top predators excrete toxins more efficiently due to their advanced physiology.
Answer: C) $5 \text{ ppm}$; Top predators accumulate toxins through biomagnification, as they consume large quantities of contaminated prey over their lifespan.
Compare the Net Primary Productivity (NPP) of a tropical rainforest to that of a desert biome. Explain the primary climatic factors responsible for this difference and how they directly influence the rate of photosynthesis in these two biomes.
- A) Desert NPP is higher due to extreme temperatures; high temperatures increase enzyme activity for photosynthesis.
- B) Tropical rainforest NPP is higher due to abundant sunlight and high precipitation; these conditions optimize water availability and light for photosynthesis.
- C) Desert NPP is higher due to low competition for resources; less competition allows individual plants to photosynthesize more efficiently.
- D) Tropical rainforest NPP is lower due to constant cloud cover reducing light; high humidity inhibits $CO_2$ uptake.
Answer: B) Tropical rainforest NPP is higher due to abundant sunlight and high precipitation; these conditions optimize water availability and light for photosynthesis.
The human population has experienced significant growth, leading to discussions about Earth's carrying capacity for humans. How does the concept of carrying capacity apply to the human population, and what role do sustainable resource use and the ecological footprint play in determining our long-term ability to thrive?
- A) Human carrying capacity is fixed and determined solely by food production; sustainable resource use is irrelevant as technology will always overcome limits.
- B) Human carrying capacity is dynamic, influenced by technology and resource consumption patterns; sustainable resource use aims to reduce the ecological footprint, allowing more people to thrive within Earth's regenerative capacity.
- C) Human carrying capacity is irrelevant due to continuous exponential growth; ecological footprint measures only carbon emissions, not overall resource use.
- D) Human carrying capacity is determined by the number of individuals, not their resource use; sustainable resource use is a concept only applicable to non-human populations.
Answer: B) Human carrying capacity is dynamic, influenced by technology and resource consumption patterns; sustainable resource use aims to reduce the ecological footprint, allowing more people to thrive within Earth's regenerative capacity.
Biodiversity hotspots are crucial for global conservation. Explain why these areas are prioritized for conservation efforts, and how habitat fragmentation, a prevalent form of habitat destruction, specifically threatens the long-term viability of species, including potential keystone species, within these hotspots.
- A) Hotspots have low species richness but high relative abundance; fragmentation increases gene flow between isolated populations.
- B) Hotspots possess high species richness and endemism but face significant threat; fragmentation reduces habitat size, increases edge effects, and isolates populations, making them more vulnerable to extinction and disrupting keystone species' roles.
- C) Hotspots are primarily marine environments with stable climates; fragmentation only affects terrestrial species, not aquatic ones.
- D) Hotspots are areas where human population density is low; fragmentation leads to increased resource availability for remaining species.
Answer: B) Hotspots possess high species richness and endemism but face significant threat; fragmentation reduces habitat size, increases edge effects, and isolates populations, making them more vulnerable to extinction and disrupting keystone species' roles.
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